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Authors
Boottanun, Patcharaporn https://orcid.org/0000-0001-5271-3094
Nagai-Okatani, Chiaki https://orcid.org/0000-0003-4084-3128
Kawanishi, Kunio https://orcid.org/0000-0003-3502-3467
Baba, Masaki https://orcid.org/0009-0002-9566-184X
Nakatsukasa, Tomofumi https://orcid.org/0000-0002-8897-4189
Ishizu, Tomoko https://orcid.org/0000-0003-3794-899X
Angata, Kiyohiko https://orcid.org/0000-0001-5561-1368
Kuno, Atsushi https://orcid.org/0000-0002-6147-6171
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Funding
Funding for this research was provided by:
Japan Society for the Promotion of Science (JP20K17136)
Japan Society for the Promotion of Science (JP24K02257)
Japan Society for the Promotion of Science (JP24K11257)
Japan Society for the Promotion of Science (JP24K19073)
Naito Foundation
National Institute of Advanced Industrial Science and Technology
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Article History
Received: 15 December 2025
Revised: 9 March 2026
Accepted: 7 April 2026
First Online: 27 April 2026
Declarations
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: All applicable international and institutional guidelines for animal care and use were followed strictly. All animals were treated according to an experimental protocol approved by the Institutional Animal Care and Use Committee of the University of Tsukuba, in compliance with the Guide for the Care and Use of Laboratory Animals (reference number: 20-146). All biological samples were obtained from Dahl-Iwai S rats (Japan SLC, Shizuoka, Japan), maintained and provided by the University of Tsukuba as part of previously approved animal studies [ ].
: The authors declare no conflict of interest.
