Handley, Jonathan M. https://orcid.org/0000-0001-6468-338X
Wilson, Joshua C. https://orcid.org/0009-0004-2472-5521
Ratcliffe, Norman https://orcid.org/0000-0002-3375-2431
Baylis, Alastair M. M. https://orcid.org/0000-0002-5167-0472
Crofts, Sarah
Pistorius, Pierre A. https://orcid.org/0000-0001-6561-7069
Funding for this research was provided by:
Rufford Foundation (Rufford Foundation)
John Cheek Trust (John Cheek Trust)
Falkland Islands Environmental Planning Department (Falkland Islands Environmental Planning Department)
Nelson Mandela University (Nelson Mandela University)
National Research Foundation (National Research Foundation)
Natural Environment Research Council (Natural Environment Research Council)
Article History
Received: 23 March 2026
Accepted: 22 June 2026
First Online: 15 July 2026
Declarations
Conflict of interest: The authors have no relevant financial or non-financial interests to disclose.
Ethics approval: Animal ethics approval was granted by Nelson Mandela University ethics committee (SCI-ZOO-014).