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Article History

Received: 21 March 2016

Accepted: 22 May 2016

First Online: 27 May 2016

Compliance with ethical standards

: This work was supported by the Defense Threat Reduction Agency under the US Army Medical Research Institute of Infectious Diseases project number 31741630.

: The content of this publication does not necessarily reflect the views or policies of the US Department of the Army, the US Department of Defense, and the US Department of Health and Human Services or of the institutions and companies with which the authors are affiliated.

: All procedures performed in studies involving animals were in accordance with the ethical standards of the institution or practice at which the studies were conducted. Research was conducted under an IACUC approved protocol in compliance with the Animal Welfare Act, PHS Policy, and other Federal statutes and regulations relating to animals and experiments involving animals. The facility where this research was conducted is accredited by the Association for Assessment and Accreditation of Laboratory Animal Care, International and adheres to principles stated in the Guide for the Care and Use of Laboratory Animals, National Research Council, 2011.

: This article does not contain any studies with human participants performed by any of the authors.

: The authors declare that they have no conflict of interest.