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Article History

Received: 8 May 2018

Revised: 20 September 2018

Accepted: 25 September 2018

First Online: 27 October 2018

Compliance with ethical standards

:

: I certify that this manuscript is original and has not been published or submitted elsewhere for publication while being considered by Applied Microbiology and Biotechnology. This study is not split up into several parts to increase the quantity of submissions and submitted to various journals or to one journal over time. No data have been fabricated or manipulated (including images) to support your conclusions. No data, text, or theories by others are presented as if they were our own. The submission has been received explicitly from all the co-authors. All the authors have contributed sufficiently to the scientific work and therefore share collective responsibility and accountability for the results.

: The authors declare that they have no conflict of interest.

: This article does not contain any studies with human participants or animals performed by any of the authors.