Funding for this research was provided by:
National Basic Research Program of China (973 Program) (2016YFD0501505)
Agriculture Research System of China (CARS-35)
National Natural Science Foundation of China (31602095)
Central Public Interest Scientific Institution Basal Research Fund (Y2016CG23)
Text and Data Mining valid from 2019-03-19
Received: 12 November 2018
Revised: 7 February 2019
Accepted: 8 February 2019
First Online: 19 March 2019
Compliance with ethical standards
: The authors declare that they have no conflict of interest.
: All guinea pig experiments were performed in a biosafety level 3 laboratory at the Lanzhou Veterinary Research Institute (LVRI) at the Chinese Academy of Agricultural Sciences (CAAS). This study was approved by the Institutional Animal Use and Care Committee of the CAAS. All guinea pigs used in the present study were humanely bred during the experiments and were euthanized at the end of the study.