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Authors
Yokomura, Masaru
Nagano, Seiji
Kawamoto, Hiroshi
Hirohashi, Yoshihiko
Torigoe, Toshihiko
Asakage, Takahiro
Katayama, Ryohei
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Funding
Funding for this research was provided by:
JST SPRING (JPMJSP2180)
Japan Society for the Promotion of Science (JP22K18383, JP24K02333)
Japan Agency for Medical Research and Development (JP24ama221210h0003, JP25ck0106795h0003, JP25ama221231h0002, 25ama221242h0001)
Nippon Foundation
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Article History
Received: 14 August 2025
Accepted: 17 December 2025
First Online: 3 January 2026
Declarations
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: Ryohei Katayama received research grants from Chugai, Nipon-Kayaku, TOPPAN, UBE, BML, EIKEN Chemical outside the submitted work. Other authors have no competing interests.
: All patients and healthy donors provided informed consent for the genetic and cell biological analyses, which were performed in accordance with protocol approved by the Institutional Review Boards of the JFCR and the Kyoto University.
: Informed consent was obtained from all individual participants included in the study.
: The authors affirm that human research participants provided informed consent for publication.
