Funding for this research was provided by:
the Strategic Priority Research Program of Chinese Academy of Sciences (XDA2006010202)
Second Tibetan Plateau Scientific Expedition and Research (STEP) program (2019QZKK010314)
National Natural Science Foundation of China (91537105, 91537211)
Article History
Received: 13 December 2019
Accepted: 18 August 2020
First Online: 29 August 2020