Article History
Received: 8 July 2020
Revised: 23 October 2020
Accepted: 17 November 2020
First Online: 30 November 2020
Compliance with ethical standards
:
: The authors declare that they have no conflict of interest.
: All research activities were fully integrated into the daily routine of the mandrills and required no manipulation of individuals. The mandrills were never deprived of food or water at any stage of the study. Our study did not meet the definition of an animal experiment as mentioned in Article 1 of the Dutch “Experiments on Animals Act”, due to the absence of potential discomfort for the study animals and the non-invasive character of the study. All applicable international, national, and/or institutional guidelines for the care and use of animals were followed. Thus, this study was in compliance with Dutch law. This study was approved by Artis Royal Zoo.