Funding for this research was provided by:
National Natural Science Foundation of China (31902349, 31472270)
Natural Science Foundation of Shandong Province (ZR2018MC030)
Shandong “Double Tops” Program
Article History
Received: 11 December 2019
Accepted: 5 February 2020
First Online: 22 February 2020
Compliance with Ethical Standards
: Animal experiments were approved by the Shandong Agricultural University Animal Care and Use Committee under approval number SDAUA-2015-017. Many efforts were made to reduce the suffering of Nile tilapia.
: The authors declare that they have no competing interests.