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Authors
Mao, Yanyan
Fan, Hengjian
Yan, Wenqing
Zhang, Na
Liu, Wenwen
Xiao, Yun
Han, Yuechen
Fan, Zhaomin
Wang, Haibo
Xia, Anping
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Funding
Funding for this research was provided by:
National Natural Science Foundation of China (31700894)
National Natural Science Foundation of China (82271172)
National Natural Science Foundation of China (81970873)
National Natural Science Foundation of China (82471174)
National Natural Science Foundation of China (8190093)
Major Program of National Fund of Philosophy and Social Science of China (82192861)
the Major Fundamental Research Program of the Natural Science Foundation of Shandong Province, China (ZR2021ZD40)
Taishan Scholar Foundation of Shandong Province (tsqn201909189)
Key Technology Research and Development Program of Shandong Province (2023CXPT038)
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Article History
Received: 6 May 2025
Accepted: 29 January 2026
First Online: 23 February 2026
Declarations
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: The authors declare that they have no conflict of interest.
: Animal care and experimental protocols were approved by the Animal Care Committee of Shandong University, People’s Republic of China. All animal experiments comply with the National Institute of Health’s Guide for the Care and Use of Laboratory Animals.
: DeepSeek (Version: DeepSeek-R1; Manufacturer: DeepSeek Co., Ltd.) was used for language polishing during October 2025. The authors take full responsibility for the content of this publication.
