Funding for this research was provided by:
Japan Society for the Promotion of Science (18K18342, 15H01619, 15H05309, 17H0582, LGP-U04, core-to-core CCSN)
Ministry of Education, Culture, Sports, Science and Technology (16H06283)
kyoto Universy SPIRITS
Text and Data Mining valid from 2019-11-11
Received: 15 April 2019
Accepted: 24 October 2019
First Online: 11 November 2019
Compliance with ethical standards
: All applicable international, national, and institutional guidelines for the care and use of animals were followed. The field observations complied with the guidelines for animal studies in the wild issued by the Wildlife Research Center of Kyoto University, Japan.
: The authors declared that they have no competing interests.