Secondi, Jean https://orcid.org/0000-0001-8130-1195
Percel, Gwendoline https://orcid.org/0000-0002-4579-345X
Arzel, Céline https://orcid.org/0000-0002-1866-2067
Davranche, Aurélie https://orcid.org/0000-0001-9969-1072
Farau, Sébastien https://orcid.org/0000-0003-3592-2446
Funding for this research was provided by:
Université d'Angers
Article History
Received: 21 January 2025
Revised: 12 September 2025
Accepted: 29 December 2025
First Online: 10 January 2026
Declarations
:
: All authors agreed with the manuscript content and all gave explicit consent to submit the version to be published.
: The study was carried out on private lands which access has been granted by landowners. No animals were handled and observation time was limited as much as possible to reduce disturbances.
: The authors declare no competing interests.