Guharajan, Roshan http://orcid.org/0000-0001-8124-5461
Arnold, Todd W.
Bolongon, Gilmoore
Dibden, Grace H.
Abram, Nicola K.
Teoh, Shu Woan
Magguna, Mohd Azzumar
Goossens, Benoit
Wong, Siew Te
Nathan, Senthilvel K. S. S.
Garshelis, David L.
Funding for this research was provided by:
Columbus Zoo and Aquarium
Point Defiance Zoo and Aquarium
Wildlife Reserves Singapore
Association of Zoos and Aquariums
International Association for Bear Research and Management
Sime Darby Foundation
Bell Museum of Natural History
Article History
Received: 15 March 2018
Revised: 9 July 2018
Accepted: 24 August 2018
First Online: 1 September 2018