Li, Yige https://orcid.org/0000-0003-3690-7299
Qiu, Siyu https://orcid.org/0009-0009-0886-8983
Wang, Fang https://orcid.org/0009-0001-5847-1193
Bi, Zhenghong https://orcid.org/0000-0002-7811-8012
Liu, Zhiyong https://orcid.org/0000-0002-9675-1233
He, Yingzi https://orcid.org/0000-0002-2248-2237
Li, Geng-Lin https://orcid.org/0000-0001-7053-4284
Funding for this research was provided by:
National Natural Science Foundation of China (82271158, 82301312)
National Natural Science Foundation of China (82192862, 82171141)
Ministry of Science and Technology of China (2021YFA1101302)
Article History
Received: 10 October 2025
Accepted: 8 April 2026
First Online: 24 April 2026
Declarations
:
: The authors declare no competing interests.
: Protocols regarding animal housing and handling are consistent with the National Institutes of Health Guidelines for the Care and Use of Laboratory Animals. The animal protocol was declared exempt by the Animal Care and Use Committee of Fudan University.
: Not applicable.