Momota, Kazuyuki https://orcid.org/0009-0004-4758-2260
Doi, Hiroyuki https://orcid.org/0000-0002-4531-165X
Kinoshita, Izumi
Sakai, Harumi
Obata, Hiroshi
Article History
Received: 3 June 2025
Accepted: 24 November 2025
First Online: 10 December 2025
Declarations
:
: Breeding management and observations were performed in accordance with “the rule of Animal-welfare of JAZA (Japanese Association of Zoos and Aquariums)( ) and “Ethic Guideline of JAZA”( ).
: The authors declare no competing interests.