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Authors
Minagawa, Kozue
Sato, Hajime https://orcid.org/0000-0003-3196-7778
Fujishima, Kanta https://orcid.org/0009-0002-9813-9950
Sakai, Yoichi https://orcid.org/0000-0002-9602-1601
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Funding
Funding for this research was provided by:
Japan Society for the Promotion of Science (22K06403)
Hiroshima University
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Article History
Received: 24 August 2025
Accepted: 19 April 2026
First Online: 7 May 2026
Declarations
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: All procedures performed in this study followed the Guidelines for the Proper Conduct of Animal Experiments outlined by the Hiroshima University Animal Research Committee (No. 004A200821), Guidelines for the Use of Animals in Research of the Association for the Study of Animal Behaviour and the Animal Behavior Society, Guidelines for the Use of Fishes in Research by the Ichthyological Society of Japan, and Guidelines for Ethological Studies by the Japan Ethological Society.
: The authors declare no competing interests.
