Funding for this research was provided by:
the Strategic Priority Research Program of the Chinese Academy of Sciences (XDA19040305)
the Science and Technology Major Project of Tibet (Z2016C01G01)
the National Science Foundation of China (41574036)
the Science Foundation of Tibet (2016ZR-15-76)
Article History
Received: 5 June 2018
Accepted: 30 October 2018
First Online: 1 January 2019