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Article History

Received: 17 August 2021

Revised: 19 January 2022

Accepted: 31 January 2022

First Online: 15 February 2022

Declarations

:

: The authors declare that there is no conflict of interest.

: The sampling was performed following the law “On approval of the amateur and sport fishing” in Ukraine (Cabinet of Ministers of Ukraine, ). Spined loaches are not commercially important fish species and their threatened status is considered to be least-concern (IUCN, ), so no fishing permission was required. The care and use of animals complied with the Ukrainian law “On the Protection of Animals from Brutal Treatment” (Verkhovna Rada of Ukraine, ).