Giraud, Gwennan https://orcid.org/0000-0002-6022-2663
Wandia, I. Nengah https://orcid.org/0000-0002-2865-2508
Hambuckers, Alain https://orcid.org/0000-0003-2594-2064
de Thier Nagelmackers, Fanny
Deleuze, Stefan https://orcid.org/0000-0003-3076-5779
Huynen, Marie-Claude
Brotcorne, Fany https://orcid.org/0000-0003-2780-3631
Funding for this research was provided by:
Fonds pour la Formation à la Recherche dans l’Industrie et dans l’Agriculture (FC031671)
Leopold III Fund for Nature Exploration and Conservation
Université de Liège (UR SPHERES, Camille Hela, Herman-van Beneden, Marie-Louise Léonard, Rolande Maréchal, Denise Dounan, Docquier)
Max Planck Institute for Evolutionary Anthropology
Article History
Received: 4 March 2025
Accepted: 8 September 2025
First Online: 7 November 2025
Declarations
:
: All authors and any responsible authorities at the organization where the work was carried out have consented to manuscript submission before the work is submitted.
: The author list includes contributors from the location where they conducted the research, who helped collect and interpret the data, and manage the long-term project.
: Not applicable.