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Article History

Received: 30 July 2018

Accepted: 23 May 2019

First Online: 30 May 2019

Compliance with ethical standards

:

: The authors declare that there are no conflicts of interest.

: This article does not contain any studies with human participants performed by any of the authors.

: The paper has been prepared and submitted in accordance with the ethical standards of the Committee on Publication Ethics (COPE). This research involved contact with wild animals (bumblebees), and we performed only observation the individual of bumblebee without capturing in consideration of influence on populations, so no ethical permit was required.