Funding for this research was provided by:
National Key R&D Program of China (No. 2024YFD2401105)
the National Natural Science Foundation of China (No. 32201108)
the Science and Technology Innovation Program of Hunan Province (No. 2022RC1167)
Hunan Provincial Natural Science Foundation of China (No. 2024JJ5251)
Hunan Provincial Natural Science Foundation of China (No. 2024JJ5053)
the Research Foundation of Education Bureau of Hunan Province (No. 24A0048)
the Research Foundation of Education Bureau of Hunan Province (No. 24B0893)
Article History
Received: 27 June 2025
Accepted: 4 March 2026
First Online: 16 March 2026
Declarations
:
: The authors declare no competing financial interest.
: All animal experiments were conducted in strict compliance with institutional animal welfare guidelines and approved by the Animal Ethics Committee of Hunan Normal University (Approval No. 2024–319).