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Article History

Received: 31 March 2020

Accepted: 24 November 2020

First Online: 4 December 2020

Compliance with ethical standards

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: The authors declare that they have no conflict of interest.

: Ethical approval was obtained from the Animal Research Ethics Committee, University of the Free State, South Africa (Student Project Number: UFS-AED2018/0040) and the South African National Biodiversity Institute’s Research Ethics and Scientific Committee (Reference: SANBI/RES/P18/20). The ethics committees use the following guiding documents to approve projects: SANS10386:2008 for the care and use of animals for scientific purposes, SANS 10379:2005 Zoo and aquarium practice, WAZA Code of Ethics and Animal Welfare, 2003, Veterinary and Para-Veterinary Professions Amendment Act, No. 19 of 1982, Professional Code of Ethics of the African Association of Zoos and Aquaria 2007 and the National Research Foundation Act 1998, Additional permitting for research use of the samples was approved under a Section 20 permit from the Department of Agriculture, Forestry and Fisheries, South Africa (Reference: 12/11/1/1/18) that permits research on animals under Section 20 of the Animal Diseases Act, 1984 (Act No 35 of 1984).

: All authors consent to publication.