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Authors
Nakao, Satoko
Sivakumar, Thillaiampalam
Ohno, Akiko
Takakuwa, Yuko
Suzuki, Hajime
Nakamura, Keiko
Ohta, Keiichiro
Tsuha, Osamu
Umemiya-Shirafuji, Rika
Yokoyama, Naoaki
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Article History
Received: 23 July 2025
Accepted: 27 November 2025
First Online: 12 December 2025
Declarations
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: All experiments were conducted in accordance with the Fundamental Guidelines for Proper Conduct of Animal Experiment and Related Activities in Academic Research Institutions under the jurisdiction of the Ministry of Education, Culture, Sports, Science and Technology. The Animal Care and Use Committee of the Obihiro University of Agriculture and Veterinary Medicine, Hokkaido, Japan approved all animal protocols (approval number: 25–17).
: The authors have no relevant financial or non-financial interests to disclose.
