Ariyama, Naomi
Hiono, Takahiro
Morita, Tatsuru
Kobayashi, Daiki
Han, Sehui
Isoda, Norikazu
Olivares, Carlos A. Flores
Ruíz, Juan Pablo
Cerda, Gerardo
Marambio, Sofía
Zamorano, Michel
Sakoda, Yoshihiro
Neira, Víctor
Funding for this research was provided by:
ANID Programa Beca Doctorado Nacional (1221863/2022)
Japan Society for the Promotion of Science (24K09260)
Takeda Science Foundation
Proyecto FII-DOC Universidad del Alba (030004I05)
Ministry of the Environment, Government of Japan (JPMEERF20254004)
Japan Agency for Medical Research and Development (JP223fa627005)
Animal Virology Laboratory, Faculty of Veterinary and Animal Sciences, University of Chile
Programa Fondecyt (1211517)
INACH Regular (RT 08–21)
Proyecto Anillo (ATE250053)
Article History
Received: 16 January 2026
Accepted: 28 May 2026
First Online: 3 June 2026
Declarations
:
: Processing of animal-derived specimens in this study was approved by the Comité Institucional de Cuidado y Uso de Animales (CICUA), University of Chile (approval number: 24760-VET-UCH). All trade and collection of animal-derived samples complied with the Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES). Specimens of Phocoena spinipinnis (CITES Appendix II) were transported under export permit 25CL000042WS. All experiments involving genetically modified organisms were authorized by the Safety Committee on Genetic Recombination Experiments of Hokkaido University (approval number: 2021-026).
: Not applicable.
: Not applicable.
: The authors declare no competing interests.