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Authors
Strong, Randy
Nelson, James F. https://orcid.org/0000-0002-7439-3033
Bogue, Molly A.
Colca, Jerry R.
Denzel, Martin
Diaz, Vivian
Finck, Brian N.
Gladyshev, Vadim N.
Horvath, Steve
Jiang, Nisi
Keller, Tracy
Kletzien, Rolf F.
Korstanje, Ron
Kumar, Navasuja
Leeuwenburgh, Christiaan
Fernandez, Elizabeth
Galecki, Andrzej
Ginsburg, Brett
Han, Melissa
Kaczorowski, Catherine
Leiser, Scott
Lopez-Cruzan, Marisa
Raj, Ken
Reifsnyder, Peter C.
Rosenthal, Nadia A.
Rosi, Susanna
Shindyapina, Anastasia
Stacpoole, Peter
Salmon, Adam B.
Tyshkovskiy, Alexander
Walter, Peter
Whitman, Malcolm
Miller, Richard A.
Harrison, David E.
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Funding
Funding for this research was provided by:
National Institute on Aging (AG022308)
National Institute on Aging (AG0222303)
National Institute on Aging (AG066346)
National Institute on Aging (AG0222307)
National Institute on Aging (AG024824)
Division of Cancer Epidemiology and Genetics, National Cancer Institute (CA034196)
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Article History
Received: 4 July 2025
Accepted: 28 August 2025
First Online: 19 September 2025
Declarations
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: The authors declare no competing interests. Note: SH has the following affiliations, but all are related to methylation data, of which there is none in this report: The Regents of the University of California are the sole owner of patents and patent applications directed at epigenetic biomarkers for which Steve Horvath is a named inventor; SH is a founder and paid consultant of the non-profit Epigenetic Clock Development Foundation that licenses these patents. SH is a Principal Investigator at the Altos Labs, Cambridge Institute of Science, a biomedical company that works on rejuvenation.
