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Article History

Received: 17 February 2025

Accepted: 29 July 2025

First Online: 14 October 2025

Change Date: 26 January 2026

Change Type: Update

Change Details: The References and citation has been corrected

Declarations

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: We declare that we have no competing interests.

: Paul gained his doctorate in mathematical biology under the supervision of Professor J.D. Murray. As a theoretician, Paul felt frustrated that model predictions were rarely tested so he trained as an experimentalist under the guidance of Professor Scott Fraser. This gave him the powerful set of combined skills in being able to determine which biological questions were amenable to mathematical modelling and which were not. Thus began a fruitful collaboration between us and the Kulesa laboratory that spanned over a decade. During this time, we combined theory with experiment to unearth a number of key findings in neural crest biology. Paul was an excellent scientist, always full of new ideas and very passionate about combining mathematics with biology to lead to insights that could not be achieved by either discipline alone. He was also very warm, kind and supportive, with a cheeky sense of humour that always made us laugh. He leaves behind his wife, Jennifer Kasemeier-Kulesa, who is also a Senior Researcher in his lab, and five children. Paul will be sorely missed by all who knew him, but he leaves behind a rich legacy of scientific work that will live long into the future.