Funding for this research was provided by:
the Second Tibetan Plateau Scientifc Expedition and Research Project (2021QZKK0301)
the National Natural Science Foundation of China (42163007)
the National Natural Science Foundation of China (41373049)
the National Natural Science Foundation of China (42003036)
the XingDian Youth Talent Plan of Yunnan Province (Grant No. XDYC-QNRC-2022-0136)
Article History
Received: 24 March 2025
Revised: 26 August 2025
Accepted: 30 November 2025
First Online: 20 December 2025