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Authors
Reimer, James Davis https://orcid.org/0000-0003-0453-8804
Kise, Hiroki https://orcid.org/0000-0002-4099-6469
Kondo, Kenta C. F.
Shimoji, Kiko
Tsuchida, Shinji https://orcid.org/0000-0002-8600-2386
Fujiwara, Yoshihiro https://orcid.org/0000-0002-1833-1866
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Article History
Received: 5 October 2025
Revised: 26 January 2026
Accepted: 6 February 2026
First Online: 9 April 2026
Declarations
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: The authors declare no competing interests.
: Not applicable.
: All necessary permits for sampling and observational field studies have been obtained by the authors from the competent authorities. The study is compliant with the CBD and Nagoya protocols.
: Genetic data available from GenBank (Accession numbers PX980026 and PX983615-PX983617). Specimens have been deposited in museums as detailed in the manuscript.
: JDR, HK, and YF conceived and designed research. JDR, HK, ST, and YF conducted sampling. HK, KCFK, and KS conducted experiments. JDR, HK, and YF contributed reagents and analytical tools. JDR, HK, KCFK, and KS analyzed data. JDR and HK wrote the manuscript. All authors read and approved the manuscript.
