Funding for this research was provided by:
Guangdong Science and Technology Department (2015A020216015)
Natural Science Foundation of Guangdong Province (2018A0303130156)
Innovative Development Fund projects of Innovation Research Institute on the South China Sea Ecological and Environmental Engineering, Chinese Academy of Sciences (ISEE2018PY02)
Article History
Received: 21 July 2019
Revised: 22 November 2019
Accepted: 31 December 2019
First Online: 27 January 2020