Kilshaw, Kerry
Montgomery, Robert A.
Campbell, Ruairidh D.
Hetherington, David A.
Johnson, Paul J.
Kitchener, Andrew C.
Macdonald, David W.
Millspaugh, Joshua J.
Funding for this research was provided by:
Recanti-Kaplan Foundation
the European Nature Trust
the Peoples Trust for Endangered Species
the Aspinall Foundation
the Whitely Animal Protection Trust
Scottish Natural Heritage
the Royal Zoological Society of Scotland