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Article History

Received: 15 September 2025

Accepted: 10 February 2026

First Online: 15 February 2026

Declarations

:

: was obtained from the Ethics Review Board of the Department of Biology, College of Natural Sciences, Jimma University (Protocol Code: Biol 602/08/2015 E.C., approved on 02/08/2015). The study was conducted in accordance with the ethical guidelines for research involving human participants established by Jimma University.

: This study involved human participants and was conducted in compliance with relevant ethical standards and institutional regulations.

: Not applicable, as this study does not include any information or images that could lead to the identification of study participants.

: Informed consent was obtained from all participants prior to their participation in the study. Participants were adequately informed about the purpose of the study, research procedures, potential risks and benefits, and their right to withdraw from the study at any time without penalty.

: The authors declare no competing interests.