Funding for this research was provided by:
Wellcome Trust (088613/Z/09/Z)
Received: 18 January 2018
Accepted: 21 March 2018
First Online: 7 May 2018
Change Date: 8 June 2018
Change Type: Correction
Change Details: This article was originally published under a CC BY-NC 4.0 license, but has now been made available under a CC BY 4.0 license. The PDF and HTML versions of the paper have been modified accordingly.
Compliance with ethical standards
: The authors declare that they have no conflict of interest.