Funding for this research was provided by:
Japan Society for the Promotion of Science (P14394, 26450204)
Received: 26 May 2016
Accepted: 8 July 2016
First Online: 26 July 2016
Compliance with Ethical Standards
: The authors declare that they have no conflict of interest.
: The nucleotide sequence data reported is available in the EMBL database under the accession number ERS1151563.