Funding for this research was provided by:
national natural science foundation of china (41871066, 41471055)
strategic priority research program (a) of the chinese academy of sciences (XDA20050101)
second tibetan plateau scientific expedition and research program (2019QZKK0503)
Received: 26 June 2021
Accepted: 11 October 2021
First Online: 25 October 2021
: The authors declare no competing interests.