Funding for this research was provided by:
JSPS KAKENHI (25304017, 23570033, 4501)
SNSF (31003A_166470)
Article History
Received: 28 June 2018
Revised: 14 August 2018
Accepted: 15 August 2018
First Online: 24 August 2018
Compliance with ethical standards
:
: The authors declare that they have no conflict of interest.
: The study complies with the current laws of the Republic of Zambia and was carried out in agreement with the Department of Fisheries: Ministry of Agriculture and Cooperatives, following the Zambian prevention of cruelty to animals act. There was no further ethical approval required. Data collection followed the ASAB/ABS (2012) guidelines for the treatment of animals in behavioural research and teaching.