Funding for this research was provided by:
Natural Environment Research Council (Case studentship 1507930)
Natural Environment Research Council (NERC CDT studentship NE/M00578X/1)
Heriot-Watt University (James Watt Scholarship)
Scottish Natural Heritage (Case studentship 1507930)
Article History
Received: 7 August 2017
Accepted: 3 May 2018
First Online: 17 June 2018
Compliance with ethical standards
:
: The authors declare that they have no conflict ofinterest.