Funding for this research was provided by:
Biotechnology and Biological Sciences Research Council (BB/M013944/1)
Text and Data Mining valid from 2019-02-14
Received: 12 November 2018
Accepted: 21 January 2019
First Online: 14 February 2019
Compliance with ethical standards
: The authors declare no conflict of interests.
: All research was carried out in accordance with the University of St Andrews Animal Welfare and Ethics Committee.