Funding for this research was provided by:
Japan Society for the Promotion of Science (16J08691, 17J08974, 16J1034, 13J05327, 25118003, 16H06301, 16H01505, 16H01726, 15K12047, 26119514, 25240020, 25118002)
Received: 21 March 2020
Revised: 8 July 2020
Accepted: 18 July 2020
First Online: 23 July 2020
Compliance with ethical standards
: We have no competing interests.
: This experiment adhered to the ethical guidelines of Kyoto University, and was approved by the Animal Experiments Committee of the Graduate School of Letters, Kyoto University. We recruited dogs and cats through a personal acquaintance network. Informed consent was obtained from all owners before their animals were tested.