Goldsborough, Zoë http://orcid.org/0000-0001-8198-5742
van Leeuwen, Edwin J. C.
Kolff, Kayla W. T.
de Waal, Frans B. M.
Webb, Christine E.
Article History
Received: 30 January 2019
Accepted: 24 August 2019
First Online: 4 September 2019
Compliance with ethical standards
:
: The authors declare that they have no conflicts of interest.
: This study involved non-human animals but was purely observational in nature, meaning that no invasive procedures were conducted whatsoever. Permission to conduct this study was granted by Royal Burgers’ Zoo Arnhem.