Funding for this research was provided by:
the National Key Research and Development Program of China (2016YFD0300106, 2017YFD0301001)
the National Natural Science Foundation of China (31771713, 31371576)
the Shandong Modern Agricultural Technology & Industry System (SDAIT-02-08)
Article History
Received: 25 June 2019
Accepted: 16 March 2020
First Online: 23 March 2020