Article History
Received: 25 April 2019
Accepted: 28 June 2019
First Online: 6 July 2019
Compliance with ethical standards
: All animal experiments comply with the ARRIVE guidelines and were carried out in accordance with the UK Animals (Scientific Procedures) Act, 1986 and associated guidelines, EU Directive 2010/63/EU for animal experiments.
: The authors declare that they have no competing interests.