Funding for this research was provided by:
National Natural Science Foundation of China (81373398, 81102422, 81501202, 81570522)
Hubei natural science foundation (2018CFB301)
Received: 18 February 2019
Accepted: 1 October 2019
First Online: 20 December 2019
Compliance with Ethical Standards
: The authors declare that there are no conflicts of interest regarding the publication of this paper.
: This article does not contain any studies with human participants performed by any of the authors.All experiments were carried out according to the National Institutes of Health Guide for the Care and Use of Laboratory Animals (publication no. 85–23, revised 1985) and the Guidelines for the Care and Use of Animals in Neuroscience Research by the Society for Neuroscience and approved by IACUC (Institutional Animal Care and Use Committee of Nanjing Medical University, NO: 14030126).