Funding for this research was provided by:
National Natural Science Foundation of China (61401271, 81401395, 81201156, 61473190, 81471733)
National Institutes of Health (EB022880)
National Key Technology R&D Program of China (2014BAI04B05)
First Online: 28 August 2018
Compliance with ethical standards
: The authors report no conflict of interest concerning the materials or methods used in this study or the findings specified in this paper.
: All procedures performed in studies involving human participants were in accordance with the ethical standards of the Huashan Institutional Review Board and with the 1964 Helsinki declaration and its later amendments or comparable ethical standards.
: Informed consent was obtained from all individual participants included in the study.