Funding for this research was provided by:
Thailand Research Fund (RSA 6080087)
Conservation Diver (n/a)
TASCMAR EU Horizon 2020 (n/a)
100th Anniversary Chulalongkorn University Fund for Doctoral Scholarship (n/a)
90th Anniversary of Chulalongkorn University Fund (n/a)
NRCT-JSPS Asian CORE COMSEA Program (n/a)
Received: 27 August 2019
Revised: 17 February 2020
Accepted: 21 February 2020
First Online: 8 May 2020
Compliance with ethical standards
: The authors declare that they have no conflict of interest.
: All applicable international, national, and/or institutional guidelines for the care and use of animals were followed by the authors.
: All necessary permits for sampling and observational field studies have been obtained by the authors from the competent authorities, through Chulalongkorn University, Thailand. The study is compliant with CBD and Nagoya protocols.
: The molecular data generated during and/or analyzed during the current study are available in the GenBank repository and can be found using accession numbers provided in Table InternalRef removed. Specimens used in the present study are deposited in the following museum: MNHN (IM-2000-28508).
: RM and MC conceived and designed the study. RM and SC conducted samplings. RM and SA conducted in-field ecological data collection. MC performed anatomical studies. RM and AW did the molecular analysis. All authors analyzed data. RM wrote the first draft of the manuscript. All authors made contributions and approved the final manuscript.