Moussy, C
Atterby, H
Griffiths, A G F
Allnutt, T R
Mathews, F
Smith, G C
Aegerter, J N
Bearhop, S
Hosken, D J
Article History
Received: 7 August 2014
Revised: 29 January 2015
Accepted: 12 February 2015
First Online: 1 April 2015
Competing interests
: The authors declare no conflict of interest.