Yu, Shujiao
Wang, Yanan
Zhao, Yanhong
Cai, Quanfeng
Liu, Bodian
Xing, Huiping
Li, Yuhu
Chen, Long
Jia, Zhihui
Funding for this research was provided by:
The Key Scientific Research Project at the Museum Level of Hubei Provincial Museum (25A05)
National Natural Science Foundation of China (22572112)
the Key Research and Development Program of Shaanxi Province (2021GY-172)
Article History
Received: 3 December 2025
Accepted: 10 March 2026
First Online: 25 March 2026
Competing interests
: The authors declare no competing interests.