Guo, Jiajie https://orcid.org/0000-0002-4116-6512
Qiu, Qiujing https://orcid.org/0009-0002-4180-0883
Yan, Xiaoyuan
Zhang, Zeying
Zhang, Xiyue
An, Na
Yin, Chengcheng
Yang, Di
Okamura, Hirohiko
Yoshida, Kaya
Sun, Hongchen
Qiu, Lihong
Funding for this research was provided by:
National Natural Science Foundation of China (81870752)
National Natural Science Foundation of China (82470970)
National Natural Science Foundation of China (81900992)
China Postdoctoral Science Foundation (2021MD703905)
Natural Science Foundation of Liaoning Province (2023-BS-097)
Article History
Received: 15 January 2025
Revised: 29 October 2025
Accepted: 3 November 2025
First Online: 5 February 2026
Competing interests
: The authors declare no competing interests.