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Authors
Shintani, Yusuke
Hayata-Takano, Atsuko https://orcid.org/0000-0002-4992-2845
Takasaki, Ichiro
Kurihara, Takashi https://orcid.org/0000-0003-4001-6771
Miyata, Atsuro
Yamano, Yui
Ikuta, Manato
Takeshita, Rei
Murata, Kenichiro
Oguri, Taisei
Asaka, Chiaki
Nunomura, Kazuto
Lin, Bangzhong
Nakagawa, Shinsaku
Okada, Takuya
Toyooka, Naoki
Takumi, Toru https://orcid.org/0000-0001-7153-266X
Ago, Yukio https://orcid.org/0000-0001-7562-4000
Takuma, Kazuhiro
Hashimoto, Hitoshi https://orcid.org/0000-0001-6548-4016
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Funding
Funding for this research was provided by:
MEXT | Japan Society for the Promotion of Science (JP19K07121, JP20H03429, JP23K06162, JP20H03429,, JP20H03429, JP20H03429, JP24K02185, JP20H00492, JP23H00395, JP24K22022, JP18H05416)
Nakatani Foundation for Advancement of Measuring Technologies in Biomedical Engineering
Japan Agency for Medical Research and Development (JP24ama121052, JP24ama121052, JP24ama121052, JP21dm0207117, JP24ama121052, JP24ama121054)
Takeda Science Foundation
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Article History
Received: 17 June 2024
Revised: 24 July 2025
Accepted: 27 August 2025
First Online: 4 September 2025
Ethical approval
: All methods in this study were conducted in compliance with relevant guidelines and regulations. All animal experiments were approved by the Animal Care and Use Committee of the Graduate School of Pharmaceutical Sciences, Osaka University (approval no. Douyaku R01-3-1 and R02-8-4). Every effort was made to minimize suffering and reduce the number of animals used. No human studies were performed.
: Y.S., A.H.-T, I.T., T.K., and H.H. hold patent JP7049637 and I.T., T.K., and N.T. hold patent JP7169592. The authors declare no other conflicts of interest.
