Varela-Andrés, Natalia https://orcid.org/0000-0003-3334-7521
Hernández-del Caño, Carlos https://orcid.org/0009-0003-4223-936X
Cebrián-León, Alejandro https://orcid.org/0000-0003-2779-7657
Blanco, Adrián https://orcid.org/0009-0005-7316-7156
Los Arcos-López de Pariza, Izaskun https://orcid.org/0009-0007-2491-8886
Fernández del Campo, Inés S. https://orcid.org/0000-0002-5034-7139
García-Losada, Sandra https://orcid.org/0009-0003-2013-3070
Arévalo, Juan Carlos https://orcid.org/0000-0003-1994-3095
Sánchez-Martín, Manuel https://orcid.org/0000-0001-8370-1336
Bajo-Grañeras, Raquel https://orcid.org/0000-0001-8634-0687
Martín, Ricardo https://orcid.org/0000-0002-7645-4901
Sánchez-Aguilera, Alberto
Merchán, Miguel A. https://orcid.org/0000-0002-1334-2822
Deogracias, Rubén https://orcid.org/0000-0002-9279-7867
Funding for this research was provided by:
University of Salamanca, Spain. Universidad de Salamanca. España
Spanish Ministry of Science, Innovation and Universities.
Fundación Tatiana Pérez de Guzmán el Bueno
EC | Directorate-General for Employment, Social Affairs and Inclusion | European Social Fund (PREP2023-001154)
Ministerio de Ciencia, Innovación y Universidades de España
Government of Junta de Castilla y León and Social European Fund
Ministry of Science, Innovation and Universities, Spain.
Instituto de Salud Carlos III and European Union
Article History
Received: 10 May 2025
Revised: 27 March 2026
Accepted: 30 April 2026
First Online: 18 May 2026
Competing interests
: The authors declare no conflict of interest.
: All the experimental procedures involving animal subjects were performed following the European Community guidelines (63/2010), the Spanish Royal Decree 53/2013 and the Order ECC/566/2015, of 20th March. Protocols were previously approved by the Bioethics Committee of the University of Salamanca, Spain.