Zhang, Yan https://orcid.org/0009-0003-3389-4055
Xue, Zhiwei
Zhang, Naibin
Zhu, Yuehua
Wu, YanZhao
Lv, Meilin
Zhang, Zhihan
Mu, Feiyu
Xing, Wenchen
Tang, Ziyi
Wang, Chunjie
Xue, Zhiyi
Zhou, Wenjing
Liu, Xiaofei
Li, Xingang https://orcid.org/0000-0002-0878-0211
Bjerkvig, Rolf
Huang, Bin https://orcid.org/0000-0002-0944-7648
Han, Mingzhi
Wang, Jian https://orcid.org/0000-0002-9482-5227
Wang, Donghai https://orcid.org/0000-0001-5048-5040
Funding for this research was provided by:
National Natural Science Foundation of China (82503652)
National Natural Science Foundation of China (82573856)
the China “111 Center”
the Key R&D Program of Shandong Province
Article History
Received: 25 August 2025
Revised: 15 January 2026
Accepted: 18 February 2026
First Online: 2 March 2026
Change Date: 1 April 2026
Change Type: Update
Change Details: The original online version of this article was revised: In the Fig 3 legend, in the sentence beginning 'Numbered annotations...', the values '[4, 8, 11, 12, 36, 40] ' should have read '[12, 4, 11, 39, 8, 35]'. These numbers refer to exon positions.
Change Date: 1 April 2026
Change Type: Correction
Change Details: A Correction to this paper has been published:
Change Details: https://doi.org/10.1038/s41388-026-03756-2
Competing interests
: The authors declare no competing interests.